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Yes, it is the Bananaquit, a member of the Tanager family! (With apologies to "To Tell the Truth"). This is my 4th blog post from our recent trip to Costa Rica, and this post is devoted to one avian family, the Tanager, or Thraupidae, in the order Passeriformes (songbirds). Tanagers reside in the Neotropics, from southern Mexico to the tip of South America. The International Ornithological Committee (IOC) recognizes 392 tanager species, distributed among 107 genera. Confusingly, only 149 of these species, less than half, are called "tanager". Another 108 are called "finch", with the rest having names including banaquit, honeycreeper, dacnis, saltator, bullfinch, seedeater, hemispingus, conebill, flowerpiercer and yellow cardinal! To make matters worse, of the four "tanagers" living in the U.S. (i.e., Summer, Scarlet, Western and Hepatic) none of them are in the Tanager family - they are all in the Cardinal family! Ohhh boy! Below is a family tree of the "True Tanagers". All the species that are in the tanager family are on the far left of the frame. To the right, in the blue box, are species that were in the tanager family, but because of genetic analysis have now been moved out. To the left of center is the Fringillidae family, with two genera that together hold 35 species of euphonia and cholophonia. On the right are three genera of the Cardinal family that hold all the "tanagers" that we see in the U.S. plus others that are in the neotropics. (Ref: Content for the family tree from Wikipedia, Tanager.) The upshot? True Tanagers are neotropical birds living well south of us, with a lot of them in, you guessed it, Costa Rica. So let's see what true tanagers look like, along with a review of the four species with tanager in the name that we enjoy north of the Rio Grande. Warning, there are 19 species in this post! Top up your coffee and let's go! An abbreviated Tanager Family Tree, showing to the right in the blue box those species moved to other families for genetic reasons, but often retaining "tanager" in their names. Let's start with birds in the Tanager family (Thraupidae). Crimson-collared TanagerCanon R5 MkII, RF 100-500mm at 500mm, 1/500sec., f/7.1, ISO 16000.
Canon R5 MkII, RF 100-500mm at 400mm, 1/250sec., f/7.1, ISO 10000. Above, presumably an immature bird has gathered banana fruit, but rather than eat it, passes it to an adult, perhaps mom or dad, and then waits to be fed. I find it intersting that the bird on the left could have eaten the banana without help, but passed it to the adult first, reinforcing their parent/offspring relationship. Feeding complete, the pair pose for a photo. Scarlet-rumped TanagerCanon R5 MkII, RF 100-500mm at 500mm, 1/320sec., f/8.0, ISO 6400.
Canon R5 MkII, RF 100-500mm at 500mm, 1/800sec., f/7.1, ISO 16000. The female, shown below, is a mix of gray, yellow/orange, reddish brown and black, with variations by sub-species. I find the colors and detail quite striking. Canon R5 MkII, RF 100-500mm at 500mm, 1/400sec., f/8.0, ISO 6400. Blue-gray TanagerCanon R5 MkII, RF 100-500mm at 500mm, 1/200sec., f/7.1, ISO 10000.
Canon R5 MkII, RF 100-500mm at 343mm, 1/800sec., f/7.1, ISO 5000. Canon R5 MkII, RF 100-500mm at 363mm, 1/200sec., f/7.1, ISO 10000. The Blue-gray Tanager has a gray head that fades to blue on the back and wings with intricate blue/green details. Palm TanagerCanon R5 MkII, RF 100-500mm at 300mm, 1/320sec., f/5.6, ISO 12800.
These images were captured in the rain. The bird is visibly wet, adding texture and detail to the feathers. Below we can see that portions of the wings are black, creating a straight line and then merging with the black tail feathers. Canon R5 MkII, RF 100-500mm at 500mm, 1/500sec., f/8.0, ISO 6400. Plain-colored TanagerCanon R5 MkII, RF 100-500mm at 500mm, 1/800sec., f/7.1, ISO 6400.
Below, the Plain-colored Tanager, with the neck, breast and belly, looking, well . . plain. Canon R7, RF 100-500mm at 500mm, 1/800sec., f/7.1, ISO 5000. Silver-throated TanagerCanon R5 MkII, RF 100-500mm at 500mm, 1/4000sec., f/7.1, ISO 5000.
Canon R5 MkII, RF 100-500mm at 500mm, 1/2000sec., f/7.1, ISO 3200. So far we have been looking at "true tanagers", colorful (with one exception!) birds with "tanager" in their name who are bona fide members of the tanager family. Next, let's stick with the "true tanagers" but look at just a few of the dozens that bear names that may surprise you! Green HoneycreeperCanon R5 MkII, RF 100-500mm at 500mm, 1/500sec., f/7.1, ISO 8000.
Above and below are photographs of the male with a black cap and mask, decurved bill, yellow on the lower mandible, and feathers that vary from blue (above) to green (below). The male below is visibly wet. Canon R5 MkII, RF 100-500mm at 500mm, 1/160sec., f/7.1, ISO 10000. Above and below, female Green Honeycreepers. Females can be totally green, or have a mix of green and blue feathers. Red-legged HoneycreeperCanon R5 MkII, RF 100-500mm at 300mm, 1/500sec., f/6.3, ISO 10000.
Canon R5 MkII, RF 100-500mm at 223mm, 1/500sec., f/5.0, ISO 10000. Below, a male flying into a nectar feeder, showing off brilliant yellow markings on the flight feathers. Canon R5 MkII, RF 100-500mm at 186mm, 1/400sec., f/5.0, ISO 10000. Below, a female Red-legged Honeycreeper with various shades of yellow and dark wing markings. The mature females will have red legs but duller than the males. Canon R5 MkII, RF 100-500mm at 451mm, 1/1000sec., f/6.3, ISO 6400. Below, a male faces off against either a female or perhaps an immature male with yellow plumage and bright red legs. Shining HoneycreeperCanon R5 MkII, RF 100-500mm at 500mm, 1/320sec., f/7.1, ISO 12800.
Above and below, a male Shining Honeycreeper showing feather detail on the back, tail, and head/neck. Shining Honeycreepers like nectar, and here are hanging onto the wire holding the feeders. Above and below, a female Shining Honeycreeper at a feeder, Sarapiqui rainforest. BananaquitCanon R5 MkII, RF 100-500mm at 500mm, 1/2000sec., f/7.1, ISO 20000.
Canon R5 MkII, RF 100-500mm at 128mm, 1/2000sec., f/8.0, ISO 10000. The images here were captured at Dave & Dave's Costa Rica Nature Park in the Sarapiqui rainforest on November 3rd. Blue DacnisCanon R6, RF 100-500mm at 472mm, 1/400sec., f/7.1, ISO 6400.
I was standing very close to the tree where this pair was foraging. We can see the birds peering up looking for insects on the underside of leaves. Above and below, a female Blue Dacnis. Buff-throated SaltatorCanon R5 MkII, RF 100-500mm at 500mm, 1/250sec., f/7.1, ISO 4000.
The sexes are similar in appearance. These images were captured in the rainforest on November 3rd. Well, that's it for the "true tanagers" ! Flame-colored Tanager Canon R5 MkII, RF 100-500mm at 500mm, 1/400sec., f/7.1, ISO 5000.
The image above was captured on November 7th in the cloud forest of the Talamanca Mountains in Costa Rica at ~8,000 feet. The images that follow below were captured in March of 2024 in Panama at ~3500 feet, near the continental divide. Canon R7, RF 100-500mm at 500mm, 1/800sec., f/8.0, ISO 640. Red-throated Ant-tanagerCanon R5 MkII, RF 100-500mm at 500mm, 1/250sec., f/7.1, ISO 2000.
The male, shown here, is a dusky red with a bright red throat, dark face, and a red crown, seen above and below. Hepatic TanagerCanon R6, RF 100-500mm at 500mm, 1/500sec., f/7.1, ISO 32000.
Although Hepatic Tanagers are residents of Central and South America, all these images are from SE Arizona on their northern breeding range. The image above was captured at Cave Creek Ranch in Portal, Arizona in April of 2023 (a male in breeding plumage), and the image below from Mt. Lemmon in October of 2021. Canon R6, RF 100-500mm at 500mm, 1/1000sec., f/7.1, ISO 4000, +0.67 EV. Canon R5 MkII, RF 100-500mm at 300mm, 1/3200sec., f/7.1, ISO 2000. Above, a male from Madera Canyon in January 2025, and below a female from the Southwestern Research Station in Cave Creek Canyon, Chiricahua Mountains, in May of 2018. Canon 7D MkII, EF 100-400mm with 1.4x III ext., at 560mm, 1/800sec., f/8.0, ISO 640, +0.67 EV. Scarlet TanagerCanon R6, RF 100-500mm with 1.4x ext., at 700mm, 1/4000sec., f/11, ISO 10000, +1.33 EV.
Canon 7D Mk II, EF 100-400mm with 1.4x III Ext.,at 560mm, 1/800sec., f/8.0, ISO 6400. Summer TanagerCanon R5 MkII, RF 100-500mm at 500mm, 1/4000sec., f/7.1, ISO 3200.
Canon R6, RF 100-400mm with 1.3x III ext., at 560mm, 1/2000sec., f/8.0, ISO 6400, +1.67 EV. Below, a female Summer Tanager at a feeder at the Paton Center for Hummingbirds in Patagonia, AZ, August 2025. Canon R5 MkII, RF 100-500mm at 500mm, 1/400sec., f/7.1, ISO 2500, +0.33 EV. Western TanagerCanon R5 MkII, RF 100-500mm with 1.4x Ext at 700mm, 1/1000sec., f/10, ISO 800.
Canon 20D, EF 70-300mm at 150mm, 1/250sec., f/5.6, ISO 400. Eighteen species so far, one more to go! Let's finish with probably the cutest bird in this series . . . . Yellow-throated EuphoniaCanon R5 MkII, RF 100-500mm at 500mm, 1/250sec., f/7.1, ISO 3200.
These images were captured at the Nest Nature Center in the Costa Rican foothills at ~2000 feet on November 2, 2025. The Yellow-throated Euphonia eats fruit, mostly mistletoe as well as a variety of berries. That's all for now!
Spring is coming, keep your eye out for new arrivals from the south! Happy trails!
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Fiery-throated Hummingbird, Talamanca cloud forest, Costa Rica, November 7, 2025. For this my third post on Costa Rica I decided to tally all of the hummingbirds we saw over the two weeks in November and post them here. These species are all unique to Central America (with some overlap north into Mexico or south into South America), with one local species that will look familiar to Arizonans. For anyone from the U.S., these are special birds. Our first group of three birds was spotted in the low altitude rainforest of NE Costa Rica, on the Caribbean side. We toured the area around Sarapique for several days, where the forest gets ~14 feet of rain a year! The lodge where we stayed, and the preserves we visited, all had covered walkways and platforms, keeping us and our gear dry (most of the time!). However, there was not a lot of light! White-necked JacobinCanon R5 m2, RF 100-500mm at 254mm, 1/1000 sec.,f/6.3, ISO 12,800.
Canon R5 m2, RF 100-500mm at 343mm, 1/1250 sec.,f/5.6, ISO 8000. The White-necked Jacobin is a strikingly beautiful bird. The "white neck" is really a patch at the nape, seen in the lower bird of the pair below. Most noticable in the male is the deep blue head and neck, offset by an almost pure white belly and tail. The tail feathers have a thin dark tip, creating a framing effect on the whole bird. The female has varying color, with "typical" markings of a blue-green breast scaled with white. They are aggressive toward one another, as the interaction below demonstrates. Note that the male at the top of the image is missing part of his secondary flight feathers on the right wing, not matched on the left, and likely due to an injury. He seems to fly well in spite of the defect. Canon R5 m2, RF 100-500mm at 128mm, 1/2500 sec.,f/8.0, ISO 10,000. For the photo geeks: These hummers are denizens of the rain forest, emphasis on a forest with high canopy and lots of rain! Which means fast moving birds in very little light. Low ambient light combined with the dark blue head and neck in the male with a bright white belly and tail make it difficult to get balanced exposure. Expose for the blue neck and the whites blow out, expose for the while tail, and the neck descends into darkness and noise! Bronze-tailed PlumeleteerCanon R5 m2, RF 100-500mm at 451mm, 1/1250 sec.,f/6.3, ISO 16,000.
The red feet are quite distinctive and a key field mark. Canon R5 m2, RF 100-500mm at 145mm, 1/1600 sec.,f/8.0, ISO 10,000. Rufous-tailed HummingbirdCanon R5 m2, RF 100-500mm at 200mm, 1/800 sec.,f/6.3, ISO 12,800.
The rufous tail itself has dark tips. Unlike the Bronze-tailed Plumeleteer, its feet are dark. Canon R5 m2, RF 100-500mm at 167mm, 1/2000 sec.,f/13, ISO 16,000. The next six species were spotted in the Talamanca Mountains, just over the continental divide on the Pacific side of Costa Rica. This area is high cloud forest, still wet, but cooler, with mixed clouds. White-throated Mountain GemCanon R5 m2, RF 100-500mm at 500mm, 1/1000 sec.,f/7.1, ISO 8000.
The White-throated Mountain-gem frequently forages in the canopy but nests in the understory. They are territorial at flowers and dominant over the Lesser Violetear, but subordinate to the Fiery-throated Hummingbird. Canon R5 m2, RF 100-500mm at 500mm, 1/250 sec.,f/7.1, ISO 4000. An aside on feather colors: The color of a bird's feather is produced in two very different ways, either as the result of pigments (red/yellow or black/gray to brown/buff) or as the result of the nanostructure of the feather surface. The structural color is produced by the interaction of incident light waves with the feather surface which reflect only certain colors back toward the viewer. The brilliant jewel like colors we see in hummingbirds is produced by the surface structure. Some of these colors are very directional, as in the throat of our next hummer, the Fiery-throated (Ref: David Allen Sibley, What It's Like to be a Bird, Knopf, New York, 2020, p. xi-xii). |
| The Fiery-throated Hummingbird lives in the cloud forests of Costa Rica and Panama, defending patches of flowers and generally showing dominance over other hummingbird species in its range. The Fiery-throated is covered with a mixture of blue, green, and yellow feathers, with a dark throat patch that turns brilliant red at the correct angle, due to the feather surface structure, hence the bird's name, Fiery-throated. (Ref: Birds of the World). |
Lesser Violetear
| The Lesser Violetear lives in mountain forests from Costa Rica to Boliva. See the ebird map to the right. The Lesser Violetear is glittering green overall with a patch of violet feathers on the sides of the head, hench the name "violet tear." Their overall color varies with the light, with a fascinating feather coloration on the throat that at certain angles looks like a knitted sweater. |
Purple-throated Mountain Gem
| The Purple-throated Mountain Gem sports iridescent green contour feathers with a white stripe behind the eye. The female (shown here) has pale rufous underparts. The male is primarily green with a purple gorget. They live in the cloud forests of Nicaragua, Costa Rica and Panama, feeding on nectar supplemented with insects. For a photograph of the male, which I did not capture on this trip, click this link, Purple-throated Mountain Gem. | Range map, Purple-throated Mountain Gem. Ref: eBird. |
Volcano Hummingbird
| The Volcano Hummingbird is among the smallest birds in the world, weighing in at 2-3 grams. Its Spanish name is Chispita, meaning "little spark." Above is a male with a brilliant purple gorget that varies from shades of purple to red depending on the subspecies. Below and following is a female with dark spotting on the throat. The Volcano lives in the highlands of Costa Rica and Panama, above ~6000 feet. (Ref: Birds of the World). |
| For the bird and photo geeks: The male captured above is feeding at small garden flowers typical for the species and its small size. The female below was photographed at at an excellent cafe and preserve where the highly attentive staff had picked flowers and hung them on the upper deck to attract the birds. As the nectar would deplete, the staff would spray the flowers with more sugar-water. The female Volcano below is feeding off of a large flower in large part because it was dripping with sugar! | Our location at the yellow square on the map. |
Talamanca Hummingbird
| If this bird looks familiar to birders in SE Arizona, it is for a good reason. In 2017 the Magnificent Hummingbird, who lived in a range from Colorado, New Mexico and Arizona down to Costa Rica was split into two separate species, the Rivoli's to the north, (including Mt. Lemmon), and the Talamanca to the south, in the mountains of the same name in Costa Rica and western Panama. |
Violet Sabrewing
| The Violet Sabrewing lives in mountain forests from southern Mexico south to western Panama, frequenting second growth forests, forest edges, banana plantations and flower gardens. These photographs were captured at the Nest Nature Center in Costa Rica, which is in the foothills at ~1600 feet. |
In Closing . . . . . .
Of this 360, about 50% live in the equatorial belt, from +10 degrees North (Costa Rica) to -10 degrees south (Brazil) latitude. About 50-54 species live in Costa Rica, making it a hummer hot spot with great places to visit and wonderful people to meet. So venture south for a visit, beef up your life-list, and get some great photographs! In this post we have seen only 10 species, but all of them a lot of fun! Next time I visit, I will have 44 more species to discover!
That's all for now! More coming soon.
Happy trails!
| My first post on Costa Rica was on November 20th, on the Resplendent Quetzal, titled Just in from Costa Rica: The Resplendent Quetzal! This post is on Cano Negro, a preserve at the far northern reaches of Costa Rica in low wetlands very close to the border with Nicaragua. See the map of Costa Rica to the right, with the red pin at Cano Negro. |
Sungrebe
| Sungrebes are very interesting birds.* First, they are not grebes, but finfoots, genetically closer to rails than to grebes. They are in the order Guiformes, family Heliornithidae, consisting of only three species, the African Finfoot, the Masked Finfoot (Asian, and endangered) and the Sungrebe, native to Central and South America, range map to the right. |
And, they hate the sun! So much for naming conventions! **
*Almost all of my blog references on U.S. birds refer to All About Birds, a Cornell website that is free to users in the U.S. However, birds that are not seen within U.S. borders are not in All About Birds, but are covered in detail in the academically oriented Birds of the World, another Cornell website, which requires a subscription. My references in this blog are mixed. I have tried to refer to free websites, but when detail is required, I do refer to Birds of the World.
**The name in fact comes from the its scientific genus Helionis, (Greek Helios, sun, and Ornis, bird), combined with grebe, because of their lobed feet, similar to grebes.
I feel fortunate to have captured these images on October 31st.
The male is unique among world species for having a specialized skin pocket under the wing (basically in the bird's "armpit") where the young can be carried even during swimming and flying. As the chicks grow, they may move to the dad's back, where they can ride even in dives. (Ref: Alvarez del Toro 1971, cited in Birds of the World Sungrebe. )
*Altrical refers to chicks that are born dependent, with minimal feathers and no ability to navigate or feed independently. These chicks are generally nest bound, and fed by their parents. Examples include most songbirds like the American Robin, and raptors like owls and hawks. Precocial chicks are born feathered and ready to go, with eggs hatching all at one time. Examples include Gambel's Quail, and Wood Ducks.
In the image of the male Sungrebe below, there seems to be an extra or at least a thicker set of underwing coverts at the base of the wing that might demarcate the pocket.
Anhinga
| Anhingas (the species) are members of the family Anhingidae, one of four families in the Order Suliformes, which includes frigatebirds, boobies, gannets, and cormorants. Note that the word "Anhinga" designates both the species described here as well as the genus it belongs to, which includes three other species, the African Darter, Oriental Darter, and the Australasian Darter. Anhingas, like cormorants, swim and dive to catch fish, and like cormorants have feathers that do not repel water, making diving easier, but requiring time out of water to air dry before they can fly. |
The male Anhinga has a black neck and body, with white patterns on the wings, and orange to red sprinkled on the head. The male seen in the images above and below was seen at Brazos Bend State Park in Texas in the spring of 2019.
The Anhingas shown below were all seen in the Cano Negro area on October 31st. The buff colored necks are typical of immature males or females and in the breeding season the male will sport turquoise blue markings around the eye, seen in the two images below. In Costa Rica Anhingas will breed when resources are plentiful, usually at the end of the dry season, March-April. My conclusion (possibly wrong!) is that this is a male at the end of breeding season that still has neck coloration of an immature. Or, perhaps there are subspecies color differences in this area of Costa Rica. If anyone has the answer, please let us all know in the comments section at the end of this post.
Below the Anhinga feet are evident, fully webbed, with a nail at the end of the toe to facilitate perching.
Anhingas are excellent swimmers, using their bill to spear fish.
Northern Jacana
| The Northern Jacana is in the order Charadriiformes, a large and diverse order known by the collective name "shorebirds", which includes plovers, oystercatchers, stilts, avocets, sandpipers, gulls, terns, skimmers, and jacanas! They are known for their long toes allowing them to walk on surface vegetation. This gives the appearance of walking on water, leading to the nickname "Jesus bird." |
Northern Jacanas have a polyandrous breeding system, whereby a female lives in a territory that covers the territories of 1-4 males. The males build multiple nests in their area, and the female mates with multiple males, laying eggs in sequence in the male's nests. The male incubates the eggs and cares for the hatchlings, which unlike Sungrebes are precocious, born able to swim, dive and feed. (Ref: Wikipedia)
Here we see a male with three chicks feeding in the marsh.
This behavior is similar to that of the Sungrebe, but in the case of the jacana, the chicks are precocious, able to walk and forage almost right away, so the jacana dad has less reason to have a special underwing pocket.
Southern Lapwing
| Southern Lapwings are in the plover family, living year round (non-migratory) from Central America all the way to the southern tip of South America. They live on grasslands and pastures, feeding on insects, small fish and aquatic invertebrates. Here we see a Southern Lapwing on grasslands adjacent to the river. They are striking birds, with a gray back and neck and a black chest patch that extends up the neck to the forehead. They have a long narrow crest, and like jacanas, a spur on each wing for defense. They breed during the rainy season, April to October, and within a day the chicks are out of the nest and feeding on their own. On this day we did not see any chicks. |
Black-collared Hawk
| Late afternoon on the river, we spotted this Black-collared Hawk, first perched, then in flight, probably looking for dinner. Black-collared Hawks eat fish with an occasional lizard or rodent, but not other birds. They live near or on fresh or brackish water from southern Mexico into South America. See the eBird range map to the right. Black-collared Hawks sport a black bib, rufous feathers, and a very short tail. |
Finally, for this season, the Kingfishers Three . . . .
Amazon Kingfisher
| Amazon Kingfishers are dark birds with a white collar and belly. The males sport a rufous breast, the females have a narrower green breast band. They live along lakeshores and slow moving rivers from Mexico to Argentina. See the range map to the right. Amazon Kingfishers eat fish and crustaceans, hunting from a perch over the water. They grab their prey, smack it on the perch, and swallow it head first. As we can see in the images above and below, the bill is large and spear-like. |
American Pygmy Kingfisher
| The American Pygmy Kingfisher lives in tropical lowlands along quiet streams and swampy areas of lakes. Their hunting style is to perch low and stay concealed, diving for fish, tadpoles and small frogs. These kingfishers are very small as their name suggests. Here we see a male with a dark head and wings and rufous collar, throat and flanks. The female has a dark green breastband with white tips to the feathers (Ref: Birds of the World). |
Green Kingfisher
| The Green kingfisher is slightly larger than the American Pygmy Kingfisher, but smaller than the Belted Kingfisher and the Ringed Kingfisher. The female (seen here) has a white collar, buffy thoat and breast with speckled green bands across the breast and uppermost belly. The male has a rufous breast. Like other kingfishers they hunt from a perch where they can see prey in the water. They eat primarily small fish. |
Happy trails!
| My wife and I just returned from two weeks in Costa Rica, where we toured three regions with Emily Carter and Holbrook Travel: first the wetlands of Cano Negro in the north, then the rainforest of Selva Verde further south, and finally the Talamanca Mountains in the province of San Gerado de Dota. In San Gerado de Dota we stayed at the Savegre Hotel, Natural Reserve and Spa, and on two successive mornings trekked a bit up the road to the edge of the cloud forest to look for the magnificent Resplendent Quetzal! |
| The Resplendent Quetzal is a member of the Trogan Family, the only family in the order Trogonidae. There are two recognized subspecies of the Resplendent Quetzal, Pharomachrus mocinno mocinno, and P.m. costaricensis, the former Guatemalan, the latter Costa Rican, seen here. The Guatemalan subspecies lives in southern Mexico and Guatemala, the Costa Rican in the mountains of Costa Rica and northern Panama. |
The map below shows our location in the mountains at about 7600 feet above sea level on the Pacific side of the continental divide.
Below, a "where's Waldo" photograph of a male Resplendent Quetzal perched behind the leaves. I resorted to manual focus to get the eye as sharp as possible (the autofocus options on my Canon R5 Mark II seemed to find everything but my subject!).
The back of the quetzal's head looks similar to the face and throat, a trait many birds share, and likely an adaption that confuses predators. If you think a bird is facing you, you may be less likely to go on the attack.
* An earlier version of this post incorrectly identified the long feather the male grows in breeding season as a "tail feather" when in fact it is part of the uppertail coverts, the feathers that cover the base of the tail feathers. This makes sense, in that the tail feathers are essential for flight and vary by species, depending on the the nature of their flight, and patterns of acquiring food. The tail coverts contribute to better aerodynamics, and a long decorative feather as part of breeding plumage would logically grow from this group. My thanks to my friend, ornithologist /birder/professional photographer Steve Vaughan for picking this up. Thanks Steve!
So, we decided to come back the next day for another try.
Up again at 4 am, coffee at 5 am, short drive down the road to the forest and our viewing spot, arriving at ~5:30 am. We were two days past the full moon, with our favorite planetary partner about to disappear over the western ridge of the mountain valley. It provided welcome light for our trek up the hill.
Sure enough, a female quetzal perched above us, image below. The females have coloration similar to the males, but a bit more muted, and without the long tail the male grows in breeding season.
These birds have a definite "cute factor", perhaps due to the short bill and large eyes on an open face.
This photo was captured at 5:32 am, 30 minutes before sunrise. Definitely low light photography.
Below, at 6:53 am, a young male quetzal arrives in a tree close to us. He is likely digesting an avocado.
It is difficult to estimate changes in population of this elusive bird, but they are considered "near threatened" with populations declining due to loss of habitat, the cloud forests.
For more on the Resplendent Quetzal, see eBird, or this essay from Living Bird magazine, Costa Ricans Join Hands to Cultivate A Quetzal Community, by Alan Poole.
More on Costa Rica soon!
Happy trails!
Fall at El Rio Preserve, Part II:Â Grebes and Warblers, with a little Chat on the side . . . . .
10/24/2025
Least Grebe (Caution: RARE BIRD ALERT!)
| The Least Grebe, like its cousin the Pied-billed Grebe, is a small duck-like water bird that lives on fresh and brackish water ranging from the Rio Grande Valley in Southern Texas all the way into South America. See the range map to the right. Least Grebes live year round within their range, so it is unusual to see one in Arizona, although as one might expect, eBird has sightings in Arizona, where it is considered a "causal resident", present intermittently throughout the year. This solitary Least Grebe has been sighted at El Rio Preserve repeatedly in September and October. See the eBird range map below for the Least Grebe sightings in our area. |
The two species seemed to get along, although I noticed that as they preened, they seemed to fluff up more when closer to the opposite species.
Pied-billed Grebe
| As noted above, Pied-billed Grebes are larger than the Least, and very common in SE Arizona. Above, two Pied-billed's at El Rio, and to the right the Pied-billed range map. The Pied-billed Grebe has an extensive range, living year round in most locations, but with some populations migrating to the northern U.S. and into Canada for summer breeding. |
| Pied-billed Grebes eat crustaceans, small fish, and whatever is available, including frogs! To the right we see a Pied-billed Grebe downing a frog at Sweetwater Wetlands in January of 2016. Yes, he did manage to swallow the whole frog, head first! For more, see the post "Grebe eats frog!", or "I can't believe he ate the whole thing!" |
Clark's Grebe
| This Clark's Grebe was spotted at Patagonia Lake State Park in March of 2019. The bird was on the far side of the lake with a lot of moist air between us, creating a slightly fuzzy image in spite of good focus and low ISO. Clark's Grebes live year round in Mexico, with populations migrating north to the western U.S. and into Canada. They frequent lakes and coastlines. See the range map to the right. This bird may have been migrating north for breeding at the time of the photograph. |
Eared Grebe
| The Eared Grebe has a thin bill and a bright red eye, seen here in non-breeding plumage. The grebe pictured here was spotted at White Water Draw in December 2019. Eared Grebes breed in colonies in shallow wetlands in the western U.S. and Canada. In the fall, after breeding, they head by the hundreds and thousands to salty inland waters, especially Mono Lake in California and the Great Salt Lake in Utah to stuff themselves with brine shrimp before heading farther south for the winter. This coincides with their fall molt providing the high calorie intake required for new feather growth. Range map is to the right. |
Let's Wrap-up with Three Warblers and a Chat . . . .
Common Yellowthroat
| The male Common Yellowthroat with his bright yellow duds and black mask looks like a mysterious highwayman, but rest assured, he is only after insects and spiders from the local leaves and branches! The Common Yellowthroat favors the edges of marshes and wetlands, breeding throughout the northern U.S. and Canada, wintering in Mexico with year round populations in Florida and the SE U.S. The adult male above was spotted at El Rio on September 21st at 7:30 am, with just enough morning sun to catch a light on the eye and create a subtle shadow on the breast. |
Yellow Warbler, Immature Male
| Jeff as always came through and told me that this looks like an immature Yellow Warbler. A Nashville would show more gray on the head, and the Tennesee and Orange-crowned would show an eye line. Young Yellow Warblers often have a more distinct eye ring than adults because the yellow is not as brilliant. Yellow Warblers are insectivores who winter in Central and South America and breed all the way from Mexico to Alaska, coast to coast! See the range map to the right. The birds pictured here are likely on their way south for the winter following the food! |
Yellow-rumped Warbler
| Our third warbler for the day is the Yellow-rumped Warbler, Audubon's. They have a yellow rump ("butter butt"), a yellow patch on the throat, and yellow patches ("pits") under the wings. Yellow-rumped Warblers have a wide range, living from the Pacific to the Atlantic and Caribbean, breeding as far north as Alaska, and wintering into Central America. They eat a wide variety of insects during the summer, as well as fruits during the winter. Unlike most warbers, their digestive tracts are capable of digesting fruit, which allows a wider range especially further north in the winter. |
And Finally, a Yellow-breasted Chat!
| For years the Yellow-breasted Chat was in the warbler family, Parulidae, but in 2017 it was given its own family, Icteriidae based in part on its large size and complex and varied vocalizations. Although wide-spread, it is generally only seen in the spring when singing, and is otherwise skulking in the shadows of dense thickets and therefore tough to spot. I spotted this bird briefly, and only once, at sunrise, 6:30 am, on September 21st, and in the shade! Once the sun was up, he was gone! |
More soon . . .
Happy trails!
* Here is a link to a prior post on El Rio: El Rio Preserve: Fall 2024
** September 22nd at about 11am in Tucson.
An American Kestrel Grabs Some Take-out . . .
The female kestrel above was perched on a high snag over the lake, just north of the main viewing deck near the parking lot. After I captured this image she took off to the east, to my right as I was standing, and disappeared into the wetlands. As she flew back to her perch I was able to get her in my viewfinder and shoot ~16 frames before she landed.
Below, she is in mid-air on her return flight.
* For more posts on American Kestrels see: New Year's Day 2018 at Fort Lowell Park.
In the image below the wings are up and we can see she has probably two dragonflies in her talons. Her rusty tail is in good light.
To see a kestrel lunching on a songbird at Ft. Lowell Park, go to the post Lunch time in Ft. Lowell Park. Caution: This post show a kestrel eating what is probably a Lark Sparrow piece by piece including the feet!
A Black-necked Stilt Swallows Them Whole . . . .
As wading shorebirds, Black-necked Stilts eat aquatic invertebrates, small crustaceans, amphibians, snails, and tiny fish. They also prey on larval mosquitoes, soldier flies, brine flies, caddisflies, dragonflies, mayflies, crickets, grasshoppers, beetles (including weevils), water-boatmen, crayfish, brine shrimp, tadpoles, and very small frogs and fish. (Ref: All About Birds).
Here we see a Black-necked Stilt working on a dragonfly, paring it down to size for the "big gulp."
Images above and below, lifting the prey from the water in the process of meal prep!
In this case playing with your food is not only okay, but an important part of the meal.
Below, after the second round she is finished pounding her prey and considers actually eating!
| From 2016, a Pied-billed Grebe downs a bullfrog at Sweetwater Wetlands. "Grebe eats frog!" or "I can't believe he ate the whole thing!" |
And, a Vermilion Flycatcher Ponders His Meal . . . .
More coming soon.
Happy trails!
The image above was captured in Summerhaven on August 23, 2025. We see three females or immature males creating a bit of a ruckus at a nectar feeder. Every summer the males are the first to migrate south, arriving in Summerhaven in early July, followed by females and immature males. By early September the gang has moved south into Mexico.
Below, a picture of a male Rufous captured on March 16, 2018 at Battiste Bed, Breakfast and Birds in Hereford, Arizona. Tony and Julie Battiste run a fantastic B&B with a large backyard feeding/viewing area, including a photo blind. This Rufous male was on his way north to breed, and stopped to get breakfast at their feeders.
Male Rufous are orange on the back and belly, with with a mix of white on the throat and a vivid iridescent-red gorget that varies in color and intensity depending on the angle of the light.
| In the spring Rufous Hummingbirds leave their wintering grounds to migrate north up the Pacific Coast, following the food (flowers and insects) up to the Pacific Northwest and Canada/Alaska. Males mate and take off early for the south, this time traveling down through the Rocky Mountains following meadow flowers. SE Arizona is one of their stops on the way to Mexico. The females and new offspring follow the males on the same route, arriving in Summerhaven usually in August. |
For more on feather color and the physics of structural color in Hummingbirds, see David Allen Sibley, What It's Like To Be a Bird, Alfred A. Knopf, New York, 2020, p 77.
You need a lot of calories to fly ~5,000 to 6,000 miles a year. Staking out and defending food sources along the way is essential.
Back in the day of mechanical shutters, one could not use flash, even with high tech strobes, with shutter speeds over ~1/250 second. If you could, the shutter would not synchronize with the flash burst, resulting in only portions of the frame being illuminated. This restriction made the use of fill flash, putting in just enough light to "fill in" darker portions of the photo, difficult for fast moving objects. I have used fill flash for hummingbirds at shutter speeds of 1/200 second, and if the bird was hovering in a stable position, I could get the eye and bill sharp, with blurring of the wings, which can look really cool. However, if the bird was darting here and there, which is often the case, everything gets blurred.
I recently discovered that high speed sync was introduced in 1995, but it has taken me 30 years to discover it. (Only the newest, hottest stuff on this blog!) Oh well . . .
In all fairness, it took the advent of electronic shutters ~ 7 years ago, to make HSS more practical. And for action photography, it took stacked sensors and rapid readout to minimize rolling shutter on electronic shutters, necessary for high speed subjects like race cars and hummingbirds. So, the manufacturers are now able to pair their speedlites (strobes) with their cameras to create a burst of rapid flashes that allows the photographer to use higher shutter speeds with flash and get even and predictable light on the subject, with minimal rolling shutter.
The image above, and the ones below, as well as a few in the preceding section, were all taken with a Canon R5 Mark II with a 580 EX II Speedlite set to HSS.
For a Canon R5 Mk II, first put a compatible speedlite on the camera. I have an aging 580 EX II, and it works. Turn on the speedlite. Turn on the camera, then go to the Red Menu (little camera icon) , second tab (2/9), and click on External Speedlite Control. Scroll down to Flash Function settings and click on it. You will see the screen below, left.
On many of my usual hummer shots the birds are side-lit or back-lit, creating shadows that are hard to deal with. The speedlite with the Better Beamer puts just enough light on the bird to get the colors and feather detail to "pop" and add a highlight off of the eye.
One remaining challange: Depth of field. In the image above the Canon R5 Mk II does a good job of locking onto the eye of the lower bird in the frame, but the upper bird is just out of the range of focus. I was shooting at f/8.0, and at f/11 or higher, I might have had both birds in focus. At high f-stops the depth of field increases, but there is less light hitting the sensor, so the ISO goes up. For the most part in 2025 high ISO's can be corrected for with post-production noise reduction software such as Topaz De Noise.
For a great video on autofocus in Canon cameras, especially the R5 Mk II, see Rudy Winston and canon autofocus explained.
Note: All images in this post processed in Adobe Lightroom Classic with noise reduction using Topaz DeNoise AI. Sharpening done as the last step.
Ed Erkes Nature Photography, Different Ways to Photograph Hummingbirds, Part I
Stay tuned, more coming soon!
Happy trails!
Above, a female Anna's sitting on a branch, which in fact, is her early morning breakfast buffet! Below a close up of the entrees . . . .
Below, our subject looks up, then to the right, likely looking for flying insects. Hummingbirds will "hawk," alighting from a branch to catch an insect in flight.
More on Anna's Hummingbirds and Their Dietary Needs . . .
| Anna's Hummingbirds live year round on the Pacific Coast from Baja to Canada, including southern Arizona. See the range map to the right. To see a recent post on Anna's, including feeding at flowers in Tucson, see Anna's Hummingbirds in the Valley, or click on the photograph above. Anna's Hummingbirds have been extensively researched. For an excellent list and description of all prior research, see Birds of the World (subscription required). Of interest, observational research in the 1970's showed that male Anna's spent ~10% of their daylight hours feeding on nectar, and ~1% feeding on insects. |
A separate study estimated that an Anna's requires 38 mg of protein a day, which can be met by consuming 38 "average flies", assuming each fly is 60% protein and 80% of the protein is utilized. Kind of like catching an occasional 'burger amid a steady diet of snow-cones!
It is raining on the mountain right now! Yahoo!
Happy trails!
American Goshawk
I downloaded the photographs to my computer on Monday. To confirm the species,, I took a photo of the computer screen for Merlin Bird ID (Cornell). Merlin was very persistent, telling me more than once that this was not a Cooper's Hawk, but an American Goshawk, one of three related accipiter* species in the U.S.
*Don't click on your dictionary app, I will explain "accipiter" below.
Below, the Goshawk crouches and takes off.
| SE Arizona is in the American Goshawk's territory, but we are unlikely to spot them often given their aversion for people. Mating pairs in the west will build nests high in the largest trees, usually firs or pines. Clutch size is 2-4 eggs. The American Goshawk is in the Order Accipitriformes along with ospreys, hawks, eagles, kites, and vultures. |
Accipiters (the Genus) are slender with short, broad, rounded wings and a long tail. They often ambush their prey, mainly small birds and mammals, capturing them after a short chase. The typical flight pattern is a series of flaps followed by a short glide. They are commonly found in wooded or shrubby areas. (Wikipedia).
Buteo is another Genus in the Family Accipitridae, and includes the hawks we commonly see in SE Arizona, including the Red-tailed, Zone-tailed, Swainson's, and Ferruginous.
So, Goshawks, Cooper's and Sharp-shinneds are commonly referred to as "Accipiters," and Red-tailed and Zone-tailed as "Buteos," after their respective Genera. In the field these two Genera look and fly differently, with Accipiters flapping and gliding, and Buteos commonly soaring.
| The goshawk has shorter and wider wings. These wings are good for quick maneuvering in tight spaces, making catching birds in flight easier. | The Red-tailed Hawk has longer more narrow wings (higher aspect ratio) good for efficient souring on thermals, in search of prey on the ground. |
Cooper's Hawk
| Cooper's Hawks live throughout North America and into Central America. Unlike Goshawks, Cooper's are comfortable around people, and often nest in trees in close proximity to houses. They are skillful in the air, chasing birds or making tight turns around buildings to attack a back-yard feeders, often forcing prey into windows. The photograph above was captured at Whitewater Draw in McNeal, Arizona, in December of 2015, the image below at Sweetwater Wetlands in Tucson in January of 2021. |
Sharp-shinned Hawk
| Sharp-shinned Hawks are the smallest of the accipiters we see in the U.S. Their flying and hunting behavior is similar to the Goshawk and Cooper's, employing flap and glide technique with tight turns to catch birds in flight, usually the size of an American Robin or smaller. They breed in Canada, building a broad flat nest high in conifers, and winter to the south, although there are many regions with year round populations. See the range map to the right. In SE Arizona we will see them during the winter, but being close to year round populations we may see them during the summer breeding season as well. |
The image above was captured at Agua Caliente in December of 2023.
Virginia's Warbler
| Virginia's Warblers winter in southern Mexico and breed in the forests of Arizona, New Mexico, Nevada, Utah and Colorado. They are ground nesters, favoring steep slopes below clumps of vegetation. (Ref: All About Birds) . Virginia's Warblers eat insects, favoring caterpillars, spiders, ants, weevils, stinkbugs and flying insects. In the images here we see a Virginia's Warbler looking for breakfast in the understory adjacent to Sabino Canyon Parkway in Summerhaven. |
Western Bluebird
Western Flycatcher, AKA Cordilleran . .
| In 1989, the Western Flycatcher was split into two separate species, the Pacific Slope and the Cordilleran. So, for the past 30 years we have been seeing the Cordilleran Flycatcher, considered the mountain cousin of the Pacific Slope, nesting in Summerhaven. However, in 2023, the two species were reunited (if they had a party, I was not invited!) as the Western Flycatcher. So, all you Cordilleran fans are now looking at, and loving, Western Flycatchers. Yahoo ! |
| In 2019, Dr. Charles Van Riper III moved his summer research on the Cordilleran Flycatcher (COFL) from Colorado to Mt. Lemmon. He worked with Dr. Harold F. Greeney, the two of them shown at the right taking a break from banding and tracking COFL's in Summerhaven in 2019. Unfortunately we lost Charles this past winter after a short illness. Everyone here on the mountain who were part of his "COFL Network" miss him dearly. Summer on the mountain does not seem right without him. |
Below, a platform nest built especially for the Western Flycatcher with a female sitting on the nest on July 13th. It is not clear if she is incubating eggs or not. Mid-July is a bit late for breeding, and this could be a second try for this season.
More coming soon. . . . .
Happy Trails!
Birds are here, but not in much evidence. This past week my wife and I walked Marshall Gulch and Bear Wallow*, with the greatest activity around a puddle (literally!) in a stream bed on the way from Bear Wallow to Mt. Bigelow. We then capped off June at the Meadow Trail on Sunday the 29th, a pause from writing this blog!
One of our finds (thanks to a fellow birder!) was a somewhat scruffy looking Red-breasted Nuthatch in Bear Wallow. Makes me want to go nuts over hatches . . . (groan!).
So let's take a look at Nuthatches!
*My thanks, as always, to Jeff Babson, ace naturalist, who led one of his classic birding expeditions to Bear Wallow on the 27th! Thanks Jeff!
Mixed Nuthatches: Red, White, Brown, and Oh, Yes, the Tiny Ones, Pygmy!
Red-breasted Nuthatch
| Red-breasted Nuthatches live in the north woods and western mountains, living year round or migrating short distances. They are one of what is known as irruptive species, those that move outside their usual range in great numbers in years when local food is scarce. Red-breasted Nuthatches eat insects during summer breeding, but switch to conifer seeds in the fall and winter. In years with low cone production they will move outside their usual range looking for seeds. This accounts for the very wide area of non-breeding regions on the range map to the right. |
| In May of 2023, with a lot of help from birding buddies Marty and Jim Herde, I was able to spot and capture a male Red-breasted Nuthatch digging a nest in a dead tree near the Mt Lemmon SkyCenter. The males will often dig out up to 4 nests, and let their mate pick the nest she wants! In this case, the female passed on this nest, which was then occupied by a pair of Pygmy Nuthatches a few weeks later. (More on this as the post progresses, keep reading!) |
Next, White, Brown, and Pygmy!
White-breasted NuthatchWhite-breasted Nuthatches eat mainly insects, as well as seeds and nuts. The species gets its "nuthatch" name from gathering large nuts that are too big to eat, then jamming them into tree bark, and "hatching" out the insides for consumption. They are black, gray and white, with a white breast that extends above the eye. The black cap is grayer in the female. |
Below, perhaps a male, caught in Hereford in January of 2017. Nuthatches will climb down trees head first foraging for insects. It is thought that climbing down a tree from top to bottom gives the bird a view of insects other birds may have missed when going from the base to the top (Ref: Woodland Woman).
Brown-headed NuthatchBrown-headed Nuthatches are the southern cousins of the other three, social birds traveling in noisy family groups throughout the southeast. See the range map to the right. They make their year round home in pine forests, eating insects and nesting in dead trees. Like other nuthatches, in cold months they will eat seeds mostly from pine cones. They have a brown cap, blue-gray back, and whitish underparts. |
And, Finally, Pygmy Nuthatches!
| Pygmy Nuthatches live year round in long-needled pine forests, primarily ponderosa pines, in the western U.S. They are cavity nesters, usually breeding in large groups in forests not affected by logging or dead wood removal. They are one of the few songbirds in North America that breed cooperatively, with a third of breeding pairs having 1-3 male helpers, often progeny or other relatives. During the winter, family groups will flock with other family groups, with up to 150 birds roosting together in one cavity in formations of squares, oblongs, triangles, diamonds, wedges, and tiers. |
Pygmy Nuthatch vs. Black-headed Grosbeak, "mano-a-mano."
Above, a Black-headed Grosbeak sits on a seed block along with a Pygmy Nuthatch, both are likely in their breeding territory and looking for food. Below the grosbeak makes threatening moves toward the nuthatch.
All birds are descendants of dinosaurs, and here the grosbeak is doing a very good Tyrannosaurus Rex impersonation!
Let's finish up with a burst of color . . . .
In Closing, a Western Tanager at 9,150 feet!
| This Western Tanager was greeting the morning sun on a pine branch on the Meadow Trail, close to the Sky Center on June 29th.* Western Tanagers winter in Mexico and Central America and summer on their breeding territories from the Rocky Mountains west to the Pacific Coast and north into Canada, all the way up to Alaska. SE Arizona is in the very southern portion of their breeding range, and I would guess a lot warmer than Alaska! *My thanks to Jim Herde for spotting this male. |
Most red birds get their red color from dietary plant pigments known as carotenoids. However, the Western Tanager gets its scarlet from a rare pigment called rhodoxanthin, also acquired in the diet probably from insects.
Western Tanagers live in open woods and eat predominantly insects, especially during breeding season, but are able to shift to fruit in fall and winter. They are tree nesters with the female building the nest. The male is attentive throughout the breeding cycle sometimes feeding her and sharing the responsibility for chasing away intruders.
Tanagers are limited to the Western Hemisphere. Most tanager species live in Central and South America, but five species live or wander into the U.S. In addition to the Western Tanager, we might spot the following four species in the U.S.:
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Stay tuned!
Happy trails!
Author
Henry Johnson, photographer and author of this site. For more detail, see About
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